In Travian version 4 and 4.4, your capital is indestructible even if the population of your capital decreases to zero and you can repopulate your capital in any situation but in Atergatis (like Travian 3.5) if population of your capital village reaches to zero you cannot repopulate it, and so it’s destroyed. You should keep it in mind that in this situation all your troops belong to your capital (in village, reinforced and/or walking) will be disappeared. So you better defend your capital and you better not to over use the “Evade in front of incoming attacks” option when it comes to your capital.